Research & Collections - Herpetology

MS Wildlife Studies, University of the Philippines - Los Baños, Philippines

BS Marine Biology, Mindanao State University - General Santos, Philippines

B.S. in Secondary Education major in Biology, De La Salle University – Dasmariñas

M.S. in Biology, De La Salle University

Ph.D. in Zoology, São Paulo State University, Brazil

M.S. in Zoology, National Museum, Federal University of Rio de Janeiro, Brazil

B.S. in Zoology, São Paulo State University, Brazil

B.S., Cornell University

Ph.D. in Species Diversification in Anolis Lizards , Washington University

B.A. in Zoology, University of Kansas, 1962

Ph.D. in Zoology, University of Kansas, 1968