Note: We hope to provide a more comprehensive list of our groups publications soon. For the time being, here is a selected bibliography of papers we published beween 2013 and 2016. 

In Press or Revision

Chan, K. O., L. L. Grismer, A. Zachariah, R. M. Brown, R. K. Abraham. 2016. Polyphyly of Asian tree toads, genus Pedostibes Günther, 1876 (Anura: Bufonidae), and the description of a new genus from Southeast Asia.  PLoS ONE.  In press.

Brown, R. M. In press.  Biogeography of Land Vertebrates.  In: Kliman, R., Gillespie, R. (Eds). Encyclopedia of Evolutionary Biology. Elsevier Ltd., Oxford, UK.

Wynn, A., A. C. Diesmos, and R. M. Brown. Two new species of Malayotyphlops from the northern Philippines, with redescriptions of M. luzonensis (Taylor) and M. ruber (Boettger). Journal of Herpetology.

Brown, R. M., Y.- C. Su., B. Barger, C. D. Siler, M. B. Sanguila, A. C. Diesmos, and D. C. Blackburn. 2016. Phylogeny of the island archipelago frog genus Sanguirana: another endemic Philippine radiation that diversified "Out-of-Palawan." Molecular Phylogeny and Evolution 94:531–536. In press.

Su, Y.-C., R. M. Brown, C.-P. Lin, and I.-M. Tso. 2016. Miocene-Pliocene island isolation sequence structured diversification of funnel web spiders of the Macrothele Taiwanensis species group (Hexathelidae) in the Taiwan-Ryukyu Archipelago. Journal of Biogeography. In press.

Hutter, C. R., J. A. Lyons, T. Kell, V. Liu, and J. M. Guayasamin. In press. The natural history, distribution, and conservation of the Lonely Rainfrog, Pristimantis eremitus. Herpetologica.

Hutter, C. R., S. M. Lambert, K. A. Cobb, Z. F. Andriampenomanana, and M. Vences. In press. A new species of bright-eyed treefrog (Mantellidae) from Madagascar, with comments on call evolution and patterns of syntopy in the Boophis ankaratra complex. Zootaxa.


Peterson, A. T., and T. Anamza. 2015. Ecological niches and present and historical geographic distributions of species: A 15-year review of frameworks, results, pitfalls, and promises. Folia Zoologica. 

Diesmos, A. C., J. L. Watters, N. A. Huron, D. R. Davis, A. C. Alcala, R. I. Crombie, L. E. Afuang, G. Gee-Das, R. V. Sison, M. B. Sanguila, M. L. Penrod, M. J. Labonte, C. S. Davey, E. A. Leone, M. L. Diesmos, E. Y. Sy, L. J. Welton, R. M. Brown, and C. D. Siler. 2015. Amphibians of the Philippines, part I: checklist of the species. Proceedings of the California Academy of Sciences 62:451–531.

Huron N. A., Realubit, N. D. C., Cobb, K. A., Brown, J. C., Bergmann, P., Morinaga, G., Diesmos, A. C., Diesmos, M. L., Brown, R. M., Siler C. D. 2015. Discovery of new island populations of the recently described False Geckos (Pseudogekko pungkaypinit and Pseudogekko ditoy): Conservation implications for the eastern Philippines. Herpetological Review 46:634–635.

Realubit, N. D. C., Diesmos, M. L. Lorenzo, A. II, De Layola, L. A., Diesmos, A. C., Brown, J. C., Brown, R. M.  Cobb, K. A., Bergman, P., Morinag, G., Siler, C. D,  Huron, N. A., Siler J. A.  2015.  Draco mindanensis (Mindanao flying lizard): Geographical distribution.  Herpetological Review 46: 216.

Brown, R. M., L. A. de Layola, A. Lorenzo II, M. L. L. Diesmos, and A. C. Diesmos. 2015. A new species of limestone karst inhabiting forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae: subgenus Lupacolus) from southern Luzon Island, Philippines. Zootaxa 4048: 191–210.

Duellman, W. E. 2015. Herpetology at Kansas: A Centennial History. SSAR. Ithaca, USA. 346 pages.

Duellman, W. E. 2015. Marsupial Frogs: Gastrotheca & Allied Genera. Johns Hopkins Press. Baltimore, USA. 407 pages.


Barley, A. J., P. J. Monnahan, R. Thomson, L. L Grismer, and R. M. Brown. 2015. Sun skink landscape genomics across Southeast Asia: the evolution of genetic and morphological diversity across a heterogeneous and fragmented landscape. Molecular Ecology 24:1696–1712. DOI: 10.1111/mec.13151

Brown, R. M., C. D. Siler, S. Richards, A. C. Diesmos, and D. C. Cannatella. 2015. Multilocus phylogeny and a new classification for Southeast Asian and Melanesian forest frogs (family Ceratobatrachidae). Zoological Journal of the Linnaean Society 174:130–168 [doi link]

Bursey, C. R., S. R. Goldberg, C. D. Siler and R. M. Brown. 2015. A new species of Cosmocerca (Nematoda: Cosmocercidae) and other helminths in Cyrtodactylus gubaot (Squamata: Gekkonidae) from the Philippines. Acta Parasitologica 60:675–­681 [doi link].

Davis, D. R., J. L. Watters, G. Köhler, C. Whitsett, N. A. Huron, R. M. Brown, A. C. Diesmos, and C. D. Siler. 2015. Redescription of the rare Philippine false gecko Pseudogekko brevipes (Reptilia: Squamata: Gekkonidae) and description of a new species. Zootaxa 4020:357–­374 [doi link].

Brown, R. M. 2015. A new species of stream frog (genus Hylarana) from the mountains of southern Mindanao Island, Philippines. Herpetologica 71:223­233 [doi link].

Hutter, C. R. and J. M. Guayasamin. 2015. Cryptic diversity concealed in the Andean cloud forests: two new species of rainfrogs (Pristimantis) uncovered through integrative taxonomy. Neotropical Biodiversity. [doi link]

Guayasamin, J. M., M. Rivera-Correa, A. Arteaga, J. Culebras, L. Bustamante, A. Pyron, N. Peñafiel, C. Morochz, and C. R. Hutter. 2015. Molecular phylogeny of stream treefrogs (Hylidae: Hyloscirtus bogotensis Group), with a new species from the Andes of Ecuador. Neotropical Biodiversity 1: 2–21. [doi link]

Guayasamin, J. M., T. Krynak, K. Krynak, J. Garcia, and C. R. Hutter. 2015. Phenotypic plasticity raises questions for taxonomically important traits: a remarkable new Andean rainfrog (Pristimantis) with the ability to change skin texture. Zoological Journal of the Linnaean Society 173: 913-928. [doi link]


Chan, K. O., L. L. Grismer, and R. M. Brown. 2014. Reappraisal of the Javanese bullfrog complex, Kaloula baleata (Müller, 1836) (Amphibia: Anura: Microhylidae), reveals a new species from northwestern Peninsular Malaysia. Zootaxa 3900:569–580.

Barley, A. J., A. Datta-Roy, K. P. Karanth, and R. M. Brown. 2015. Sun skink diversification across the Indian-Southeast Asian biogeographical interface.  Journal of Biogeography [doi link]

Siler, C. D. , L. J. Welton, D. R. Davis, J. L. Watters, C. S. Davey, A. C. Diesmos, M. L. Diesmos, and R. M. Brown.  2014.  Taxonomic revision of the Pseudogekko compresicorpus Complex (Reptilia: Squamata: Gekkonidae), with descriptions of three new species.  Herpetological Monographs 28:110–139.[doi link]

Welton, L. J., C. D. Siler, S. N. Travers, and R. M. Brown. 2014. Integrative taxonomy and phylogeny-based species delimitation of Philippine water monitor lizards (Varanus salvator Complex) with descriptions of two new cryptic species.  Zootaxa 3881:201–227.[doi link]

Diesmos, A. C., A. C. Alcala, C. D. Siler, and R. M. Brown. 2014. Status and Conservation of Philippine Amphibians. Pp. 310–336 In: Heatwole, H., and I. Das (Eds.). Conservation Biology of Amphibians of Asia.  Status of Conservation and Decline of Amphibians: Eastern Hemisphere. Natural History Publications (Borneo), Kota Kinabalu, Malaysia.

Duellman, W. E., A. J. Barley, and P. J. Venegas. 2014. Cryptic species diversity in marsupial frogs (Anura: Hemiphractide: Gastrotheca) in the Andes of northern Peru. Zootaxa 3768:159–177. [doi link]

Folly, M., F. Hepp, S. P. Carvalho-e-Silva, and W. E. Duellman. 2014. Taxonomic status and redescription of Flectonotus ulei Miranda-Ribeiro, 1926 (Anura: Hemiphractidae), with a key to species of Fritziana. Zoologia 31:393–399. [doi link]

Kin Onn, C., R. M. Brown, N. Ahmad and L. Lee. Grismer. 2014. A new species of frog (Amphibia: Anura: Ranidae) of the Hylarana signata Complex from Peninsular Malaysia. Herpetologica 70:228–240. [[doi link].

Richards, S. J., P. Oliver, and R. M. Brown. 2014. A new scansorial species of Platymantis Günther 1858 (Anura: Ceratobatrachidae) from Manus Island, Admiralty Archipelago, Papua New Guinea. Pp. 123–134 In: Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea Monograph Series Vol. 2, The Entomological Society of Latvia, Lettland, Latvia. Leviton, A. E., R. M. Brown, and C. D. Siler. 2014. The dangerously venomous snakes of the Philippine archipelago, with identification keys and species accounts. Pp. 473–530 In: G. C. Williams and G. C. Gosliner (eds.), The Coral Triangle: the 2011 Hearts Philippine Biodiversity Expedition. California Academy of Sciences, San Francisco, CA.

Welton, L. J., P. L. Wood Jr., J. R. Oaks, C. D. Siler, and R. M. Brown. 2014. Fossil-calibrated phylogeny and historical biogeography of Southeast Asian water monitors (Varanus salvator Complex). Molecular Phylogenetics and Evolution 74:29–37. [doi link]

Gonzales, P., Y.-C. Su, C. D. Siler, A. Barley, M. B. Sanguila, A. C. Diesmos and, R. M. Brown. 2014. Archipelago colonization by ecologically dissimilar amphibians: evaluating the expectation of common evolutionary history of geographical diffusion in co-distributed rainforest tree frogs in islands of Southeast Asia. Molecular Phylogenetics and Evolution 72: 35–41. [doi link]

Siler, C. D., A. Lira-Noriega, and R. M. Brown. 2014. Conservation genetics of Australasian sailfin lizards: flagship species threatened by coastal development and insufficient protected area coverage. Biological Conservation 169:100–108. [doi link]

Siler, C, D., A. A. Dececchi, C. L. Merkord, D. R. Davis, T. J. Christiani, and R. M. Brown. 2014. Cryptic diversity and population genetic structure in the rare, endemic, forest-obligate, geckos of the Philippines. Molecular Phylogenetics and Evolution 70:204–209. [doi link]

Brown, R. M., Jennifer A. Weghorst, Karen V. Olson, Mariano R. M. Duya, Anthony J. Barley, Liza V. Duya, Myron Shekelle, Irene Neri-Arboleda, Jacob A. Esselstyn, Nathaniel J. Dominy, Perry S. Ong, Gillian L. Moritz, Adrian Luczon, Mae L. D. Diesmos, Arvin C. Diesmos and Cameron D. Siler. 2014. Conservation genetics of the Philippine tarsier: cryptic genetic variation restructures conservation priorities for an island archipelago primate. PLoS ONE 9:e104340. [doi link]

Rocha, L. A., A. Aleixo, G. Allen, F. Almeda, C. Baldwin, M. Barclay, J. Bates, A. Bauer, F. Benzoni, C. Berns, M. Berumen, D. C. Blackburn, S. Blum, F. Bolaños, R. Bowie, R. Brit, R. M. Brown, et. al. 2014.  Specimen collection: an essential tool. Science 344:814–815. [doi link] Rocha, L. A., A. Aleixo, G. Allen, F. Almeda, C. Baldwin, M. Barclay, J. Bates, A. Bauer, F. Benzoni, C. Berns, M. Berumen, D. C. Blackburn, S. Blum, F. Bolaños, R. Bowie, R. Brit, R. M. Brown, et. al. 2014.  Specimen collection: an essential tool. Science 344:814–815. [doi link]

Siler, C. D., Linkem, C. W., Cobb, K., Watters, J. L., Cummings, S. T., Diesmos, A. C., & Brown, R. M. (2014). Taxonomic revision of the semi-aquatic skink Parvoscincus leucospilos (Reptilia: Squamata: Scincidae), with description of three new species. Zootaxa 3847(3):388-412.

Siler, C. D., Oaks, J. R., Cobb, K., Ota, H., Brown, R. M. (2014), Critically endangered island endemic or peripheral population of a widespread species? Conservation genetics of Kikuchi's gecko and the global challenge of protecting peripheral oceanic island endemic vertebrates. Diversity and Distributions, 20:756–772.[[doi link]


Hutter C.R., J. M. Guayasamin, and J. J. Wiens. 2013. Explaining Andean megadiversity: the evolutionary and ecological causes of glassfrog elevational richness patterns. Ecology Letters 16: 1135–1144. [doi link]

Gamble, T., A. J. Geneva, R. E. Glor, and D. Zarkower. In Press. Anolis sex chromosomes are derived from a single ancestral pair. Evolution. [doi link]

Schmid, M., C. Steinlein, J. P. Bogart, T. Haaf, L. Nanda, E. del Pino, W. E. Duellman, and S. B. Hedges. 2013. The hemiphractid frogs: phylogeny, embryology, life history, and cytogenetics. Cytogenetics and Genome Research 138:69–384.

Siler, C. D., A. Lira-Noriega, and R. M. Brown. 2013. Conservation genetics of Australasian Sailfin Lizards: flagship species threatened by coastal development and insufficient protected area coverage. Biological Conservation 169:100–108 [doi link]

Ng, J., A. L. Kelly, D. J. MacGuigan, and R. E. Glor. 2013. The role of heritable and dietary factors in the sexual signal of a Hispaniolan Anolis lizard, Anolis distichus. Journal of Heredity 104:862-873. [doi link]

Hutter C. R., S. Escobar-Lasso, J. A. Rojas-Morales., P. A. Gutièrrez-Càrdenas, H. Imba, and J. M. Guayasamin. 2013. High site fidelity, aggressive interactions and vocalisations of the Red-spotted Glassfrog, Nymphargus grandisonae, Cochran & Goin 1970 (Anura: Centrolenidae). The Journal of Natural History 47: 3011-3032 [doi link]

Blackburn, David C., and W. E. Duellman. 2013. Brazilian marsupial frogs are diphyletic (Anura: Hemiphractidae: Gastrotheca). Molecular Phylogenetics and Evolution 68:709–714. [doi link]

Grismer, L. L., P. L. Wood Jr., S. Anuar, M. A. Muin, E. S. H. Quah, J. A. McGuire, R. M. Brown, and N. V. Tri. 2013. Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia. Zoological Journal of the Linnaean Society 169:849–880 [doi link]

Wollenberg, K. C., I. J. Wang, R. E. Glor, and J. B. Losos (2013) Determinism in the diversification of Hispaniolan trunk-ground anols(Anolis cybotes species complex). Evolution. [doi link]

Duellman, William E. 2013. An elusive new species of marsupial frog (Anura: Hemiphractidae: Gastrotheca) from the Andes of northern Peru. Phyllomedusa, 12, 3–11.

Scantlebury, D. P. 2013. Diversification rates have declined in the Malagasy herpetofauna. Proc. Royal Soc. B. [doi link]

Brown, R. M., C D. Siler, C. H. Oliveros, J. A. Esselstyn, A. C. Diesmos, P. A. Hosner, C. W. Linkem, A. J. Barley, J. R. Oaks, M. B. Sanguila, L. J. Welton, D. S. Blackburn, R. G. Moyle, A. T. Peterson, and A. C. Alcala. 2013. Evolutionary processes of diversification in a model island archipelago. Annual Review of Ecology, Evolution, and Systematics 44:411–435 [doi link]

Ascher, J. M., A. J. Geneva, J. Ng, J. D. Wyatt, and R. E. Glor. 2013. Phylogenetic analyses of novel squamate adenovirus sequences in wild-caught Anolis lizards. PLOS ONE. [doi link]

Siler, C, D., A. A. Dececchi, C. L. Merkord, D. R. Davis, T. J. Christiani, and R. M. Brown. 2013. Cryptic diversity and population genetic structure in the rare, endemic, forest-obligate, geckos of the Philippines. Molecular Phylogenetics and Evolution 70:204–209 [doi link]

Lambert, S. M., A. J. Geneva, D. L. Mahler, and R. E. Glor. 2013. Using genomic data to revisit a classic example of reproductive character displacement in Haitian Anolis lizards. Molecular Ecology 22:3981-3995. [doi link] [see also Losos and Leal's perspective on this article]

Brown, R. M., S, J. Richards, and T. S. Broadhead. 2013. A new shrubfrog in the genus Platymantis (Ceratobatrachidae) from the Nakanai Mountains of eastern New Britain Island, Bismarck Archipelago. Zootaxa 3710: 31–45 [doi link]

Chan, K. O., D. C. Blackburn, R. W. Murphy, B. L. Stuart, D. A. Emmett, Cuc Thu Ho, and R. M. Brown. 2013. New species of narrow-mouthed frog of the genus Kaloula from eastern Indochina. Herpetologica 69:329–341 [doi link]

Algar, A. C., D. L. Mahler, R. E. Glor, and J. B. Losos (2013) Niche incumbency, dispersal limitation and climate change shpe geographical distributions in a species-rich island adaptive radiation. Global Ecology and Biogeography 22:391-402. [doi link]

Barley, A., J. White, A. C. Diesmos, and R. M. Brown. 2013. The challenge of species delimitation at the extremes: diversification without morphological change in Philippine sun skinks. Evolution 67:3556-3572 [doi link]

Hertz, P. E., Y. Arima, A. Harrison, R. B. Huey, J. B. Losos, and R. E. Glor. 2013. Tempo and bone in evolution: asynchonous trait evolution in Anolis lizards. Evolution. [doi link]

Brown, R. M., and C. D. Siler. 2013. Spotted stream frog diversification at the Australasian faunal zone interface, mainland versus island comparisons, and a test of the Philippine ‘dual-umbilicus’ hypothesis. Journal of Biogeography 41:182–195 [doi link]

Ng, J., E. L. Landeen, R. M. Logsdon, R. E. Glor. 2013. Correlation between Anolis lizard dewlap phenotype and environmental variation indicates adaptive divergence of a signal important to sexual selection and species recognition. Evolution 67:573–582. [doi link]

Linkem, C. W., and R. M. Brown. 2013. Systematic revision of the Parvoscincus decipiens (Boulenger 1894) complex of Philippine forest skinks (Squamata: Scincidae: Lygosominae) with descriptions of seven new species. Zootaxa 3700: 501–533. [doi link]

Blackburn, D. C, C. D. Siler, A. C. Diesmos, J. A. McGuire, D. C. Cannatella, and R. M. Brown. 2013. An adaptive radiation of frogs in a Southeast Asian island archipelago. Evolution 67:2631–2646 [doi link]

Welton, L. J., C. D. Siler, J. R. Oaks, A. C. Diesmos, and R. M. Brown. 2013. Multilocus phylogeny and Bayesian estimates of species boundaries reveal hidden evolutionary relationships and cryptic diversity in Southeast Asian monitor lizards. Molecular Ecology 22:3495–3510 [doi link]

Siler, C. D., L. J. Welton, A. A. Rock and R. M. Brown. 2013. Population genetic structure and revised geographic range for the tridactyl skink (Brachymeles muntingkamay) from Luzon Island, Philippines. Herpetological Review 44:30–33 [PDF link]

Brown, R. M., C. D. Siler, C. H. Oliveros, L. J. Welton, A. Rock, J. Swab, M. Van Weerd, J. van Beijnen, E. Jose, D. Rodriguez, E. Jose, and A. C. Diesmos. 2013. The amphibians and reptiles of Luzon Island, Philippines, VIII: the herpetofauna of Cagayan and Isabela Provinces, northern Sierra Madre Mountain Range. Zookeys 266:1–120 [doi link]

Linkem C. W., R. M. Brown, C. D. Siler, B. J. Evans, C. C. Austin, D. T. Iskandar, A. C. Diesmos, J. Supriatna, N. Andayani, and J. A. McGuire. 2013. Stochastic faunal exchanges drive diversification in widespread Wallacean and Pacific Island lizards (Squamata: Scincidae: Lamprolepis smaragdina). Journal of Biogeography 40:507–520 [doi link]

M Böhm, et al. 2013. The conservation status of the world’s reptiles. Biological Conservation 157:372–385 [doi link]

Siler, C. S., C. H. Oliveros, A. Santannen, and R. M. Brown. 2012. Multilocus phylogeny reveals unexpected patterns of regional and morphological diversification in Asian Wolf Snakes (genus Lycodon). Zoologica Scripta 42:262–277 [doi link]

Welton, L. J., C. D. Siler, C. W. Linkem, A. C. Diesmos, M. L. Diesmos, E. Sy, and R. M. Brown. 2013. Dragons in our midst: phyloforensics of illegally traded Southeast Asian water monitor lizards. Biological Conservation 159:7–15 [doi link]

Oaks, J., R., J. Sukumaran, J. A. Esselstyn, C. W. Linkem, C. D. Siler, M. T. Holder, and R. M. Brown. 2013. Evidence for Pleistocene-driven diversification? A caution for interpreting ABC inferences of simultaneous historical events. Evolution 67:991–1010 [doi link]