Selected Publications


Girard, M. G., Davis, M. P., Baldwin, C. C., Dettaï, A., Martin, R. P., & Smith, W. L. 2022. Molecular phylogeny of the threadfin fishes (Polynemidae) using ultraconserved elements. Journal of Fish Biology, 100 (3), 793– 810. DOI: 10.1111/jfb.14997

Chenchen Shen & Gloria Arratia. 2022. Re-description of the sexually dimorphic peltopleuriform fish Wushaichthys exquisitus (Middle Triassic, China): taxonomic implications and phylogenetic relationships, Journal of Systematic Palaeontology. 19 (19), 1317-1342. DOI: 10.1080/14772019.2022.2029595

Martin, R. P., Davis, M. P., & Smith, W. L. 2022. The impact of evolutionary trade-offs among bioluminescent organs and body shape in the deep sea: A case study on lanternfishes. Deep Sea Research Part I: Oceanographic Research Papers, 184, 103769. DOI: 10.1016/j.dsr.2022.103769


Regmi, B., Douglas, Edds, & Douglas, M.E. 2020. Geometric morphometric analyses define riverine and lacustrine species flocks of Himalayan snowtrout (Cyprinidae: Schizothorax) in Nepal. Aquatic Biology 30: 19-31. DOI: 10.3354/ab00737

Matthew G. Girard, Matthew P. Davis, W. Leo Smith. 2020. The Phylogeny of Carangiform Fishes: Morphological and Genomic Investigations of a New Fish Clade," Copeia, 108(2), 265-298. DOI: 10.1643/CI-19-320

Regmi, Binod & Douglas, Marlis & Edds, David & Wangchuk, Karma & Lu, Chang & Khanal, Gopal & Norbu, Pema & Norbu, Sangay & Dorji, Sonam & Tshering, Singye & Angel, Zachary & Chafin, Tyler & Zbinden, Zachery & Douglas, Michael. 2020. The Himalayan Uplift and the Evolution of Aquatic Biodiversity Across Asia: Snowtrout (Cyprininae: Schizothora x) as a Test Case. DOI: 10.1101/2020.10.12.336149

Alex J. Maile, Zachary A. May, Emily S. DeArmon, Rene P. Martin, Matthew P. Davis. 2020. Marine Habitat Transitions and Body-Shape Evolution in Lizardfishes and Their Allies (Aulopiformes). Copeia, 108(4), 820-832. DOI: 10.1643/CG-19-300


Arratia, Gloria and Quezada-Romegialli, Claudio. 2019. The South American and Australian percichthyids and perciliids. What is new about them?. Neotropical Ichthyology 17(1), e180102. DOI: 10.1590/1982-0224-20180102


W. Leo Smith, Chesney A. Buck, Gregory S. Ornay, Matthew P. Davis, Rene P. Martin, Sarah Z. Gibson, Matthew G. Girard. 2018. Improving Vertebrate Skeleton Images: Fluorescence and the Non-Permanent Mounting of Cleared-and-Stained Specimens," Copeia, 106(3), 427-435. DOI: 10.1643/CG-18-047

W. Leo Smith, Elizabeth Everman, Clara Richardson. 2018. Phylogeny and Taxonomy of Flatheads, Scorpionfishes, Sea Robins, and Stonefishes (Percomorpha: Scorpaeniformes) and the Evolution of the Lachrymal Saber. Copeia, 106(1), 94-119. DOI: 10.1643/CG-17-669

Martin RP, Olson EE, Girard MG, Smith WL, Davis MP. 2018. Light in the darkness: New perspective on lanternfish relationships and classification using genomic and morphological data. Mol Phylogenet Evol. 2018 121:71-85. DOI: 10.1016/j.ympev.2017.12.029

Martin, RP, Davis, MP. The evolution of specialized dentition in the deep-sea lanternfishes (Myctophiformes). Journal of orphology. 2020; 281: 536– 555. DOI: 10.1002/jmor.21120

Ghedotti MJ, Gruber JN, Barton RW, Davis MP, Smith WL. 2018. Morphology and evolution of bioluminescent organs in the glowbellies (Percomorpha: Acropomatidae) with comments on the taxonomy and phylogeny of Acropomatiformes. J Morphol. 279(11): 1640-1653. DOI: 10.1002/jmor.20894

Arratia, G. 2018. Otomorphs (= otocephalans or ostarioclupeomorphs) revisited. Neotropical Ichthyology.


Arratia G, Vila I, Lam N, Guerrero CJ, Quezada-Romegialli C. 2017. Morphological and taxonomic descriptions of a new genus and species of killifishes (Teleostei: Cyprinodontiformes) from the high Andes of northern Chile. PLoS ONE 12(8): e0181989. DOI: 10.1371/journal.pone.0181989

Arratia G, Quezada-Romegialli C. 2017. Understanding morphological variability in a taxonomic context in Chilean diplomystids (Teleostei: Siluriformes), including the description of a new species. PeerJ 5:e2991 DOI: 10.7717/peerj.2991

Gloria Arratia. 2017. New Triassic teleosts (Actinopterygii, Teleosteomorpha) from northern Italy and their phylogenetic relationships among the most basal teleosts, Journal of Vertebrate Paleontology, 37:2, e1312690, DOI: 10.1080/02724634.2017.1312690


Girard MG, Smith WL. 2016. The phylogeny of marine sculpins of the genus Icelinus with comments on the evolution and biogeography of the Pseudoblenninae. Zootaxa 4171(3): 549-561. 

W. Leo Smith, Jennifer H. Stern, Matthew G. Girard, Matthew P. Davis. 2016. Evolution of Venomous Cartilaginous and Ray-Finned Fishes, Integrative and Comparative Biology 56(5): 950–961, DOI: 10.1093/icb/icw070

Davis, M. P., Sparks, J. S., & Smith, W. L. 2016. Repeated and Widespread Evolution of Bioluminescence in Marine Fishes. PloS one, 11(6), e0155154. DOI: 10.1371/journal.pone.0155154


Wiley EO, Fuiten AM, Doosey MH, Lohman BK, Merkes C, Azuma M. 2015. The Caudal Skeleton of the Zebrafish, Danio rerio, from a Phylogenetic Perspective: A Polyural Interpretation of Homologous Structures. Copeia. 103(4):740-750. DOI: 10.1643/CG-14-105

Owens, H.L. 2015. Evolution of codfishes (Teleostei: Gadinae) in geographical and ecological space: evidence that physiological limits drove diversification of subarctic fishes. J. Biogeogr., 42: 1091-1102. DOI: 10.1111/jbi.12483

Nancy I. Holcroft, E. O. Wiley. 2015. Variation in the Posttemporal-Supracleithrum Articulation in Euteleosts. Copeia, 103(4), 751-770. DOI: 10.1643/CG-14-099

Doosey, M.H., Wiley, E.O. 2015. Epural bones in teleost fishes: a problem of phylogenetic homology. Ichthyol Res 62, 131–144. DOI: 10.1007/s10228-014-0413-z

Gloria Arratia. 2015. Complexities of Early Teleostei and the Evolution of Particular Morphological Structures through Time," Copeia, 103(4), 999-1025. DOI: 10.1643/CG-14-184


Smith WL, Busby MS. 2014. Phylogeny and taxonomy of sculpins, sandfishes, and snailfishes (Perciformes: Cottoidei) with comments on the phylogenetic significance of their early-life-history specializations. Mol Phylogenet Evol. 79:332-52. DOI: 10.1016/j.ympev.2014.06.028

Michael H. Doosey, Nicholas D. Domke. 2014. Early Development of the Caudal Fin Skeleton of Capelin, Mallotus villosus (Osmeridae). Copeia, 2014(2), 355-365. DOI: 10.1643/CG-13-098

Davis, M.P., Holcroft, N.I., Wiley, E.O., Sparks, J.S., and Smith, W.L. 2014. Species-specific bioluminescence facilitates speciation in the deep sea. Marine Biology 161: 1139-1148.

Price, S.A., Schmitz, L., Oufiero, C.E., Eytan, R.I., Dornburg, A., Smith, W.L., Friedman, M., Near, T.J., and Wainwright, P.C. 2014. Two waves of colonization straddling the K–Pg boundary formed the modern reef fish fauna. Proceedings of the Royal Society of London, B 281: 20140321

Sparks, J.S., Schelly, R.C., Smith, W.L., Davis, M.P., Tchernov, D., Pieribone, V.A., and Gruber, D.F. 2014. The covert world of fish biofluoresence: A phylogenetically widespread and phenotypically variable phenomenon. PLoS ONE 9: e83259. 

Stewart, T.A., Smith, W.L., and Coates, M.I. 2014. The origins of adipose fins: An analysis of homoplasy and the serial homology of vertebrate appendages. Proceedings of the Royal Society of London, B 281: 20133120.


Arratia, G. 2013. Morphology, taxonomy, and phylogeny of Triassic pholidophorid fishes. Memoir 13, Journal of Vertebrate Paleontology 23, Supplement to 6: 1-138.

Arratia, G., and Schultze, H.-P. 2013. Outstanding features of a new Late Jurassic pachycormiform fish from the Kimmeridgian of Brunn, Germany and comments on current understanding of pachycormiforms. Pages 87-120, In: Mesozoic Fishes 5–Global Diversity and Evolution. Arratia, G., Schultze, H.-P., and Wilson, M.V.H. (eds.). Verlag Dr. Friedrich Pfeil, München, Germany.

Betancur-R., R., Broughton, R.E. , Wiley, E.O., Carpenter, K., López, J.A., Li, C., Holcroft, N.I., Arcila, D., Sanciangco, M., Cureton, J., Zhang, F., Buser, T., Campbell, M., Rowley, T., Ballesteros, J.A., Lu, G., Grande, T., Arratia, G., and Ortí, G. 2013. The tree of life and a new classification of bony fishes. PLoSONE Currents Tree of Life. (doi:10.1371/currents.tol.53ba26640df0ccaee75bb165c8c26288)

Borden, W.C., Grande, T., and Smith, W.L. 2013. Comparative osteology and myology of the caudal fin in the Paracanthopterygii (Teleostei: Acanthomorpha). Pages 419-455, In: Mesozoic Fishes 5–Global Diversity and Evolution. Arratia, G., Schultze, H.-P., and Wilson, M.V.H. (eds.). Verlag Dr. Friedrich Pfeil, München, Germany.

Broughton, R.E., Betancur-R., R.,Li, C., Arratia, G., and Ortí, G. 2013. Multi-locus phylogenetic analysis reveals the pattern and tempo of bony fish evolution. PLoSONE Currents Tree of Life 2013, Apr 13. (doi:10.1371/currents.tol.2ca8041495ffafd0c92756e75247483e) 

Davis, M.P., Arratia, G., and Kaiser, T.M. 2013. The first Fossil Shellear (Gonorynchiformes: Kneriidae) from the Eocene lake of Mahenge (Tanzania). Pages 325-362, In: Mesozoic Fishes 5–Global Diversity and Evolution. Arratia, G., Schultze, H.-P., and Wilson, M.V.H. (eds.). Verlag Dr. Friedrich Pfeil, München, Germany.

Davis, M.P., Midford, P.E., and Maddison, W. 2013. Exploring power and parameter estimation of the BiSSE method for analyzing species diversification. BMC Evolutionary Biology. 13:38. 

Dikow, R.B. and Smith, W.L. 2013. Complete genome sequences provide a case study for the evaluation of gene-tree thinking. Cladistics 29:672-682.

Dikow, R.B. and Smith, W.L. 2013. Genome-level homology and phylogeny of Vibrionaceae (Gammaproteobacteria: Vibrionales) with three new complete genome sequences. BMC Microbiology 13:80.

Ghedotti, M.J., and Davis, M.P. 2013. Phylogeny, classification, and evolution of salinity tolerance of the North American topminnows and killifishes, Family Fundulidae (Teleostei: Cyprinodontiformes).  Fieldiana: Life and Earth Sciences. 7:1-65.

González-Rodríguez, K. A., Schultze, H.-P., and Arratia, G. 2013. Miniature armored teleosts from the Albian-Cenomanian (Cretaceous) of Mexico. Pages 457-487, In: Mesozoic Fishes 5–Global Diversity and Evolution. Arratia, G., Schultze, H.-P., and Wilson, M.V.H. (eds.). Verlag Dr. Friedrich Pfeil, München, Germany.

Grande, T., Borden, W.C., and Smith, W.L. 2013. Limits and relationships of Paracanthopterygii: A molecular framework for evaluating past morphological hypotheses. Pages 385-418, In: Mesozoic Fishes 5–Global Diversity and Evolution. Arratia, G., Schultze, H.-P., and Wilson, M.V.H. (eds.). Verlag Dr. Friedrich Pfeil, München, Germany.

McMahan, C.D., Chakrabarty, P., Sparks, J.S., Smith, W.L., and Davis, M.P. 2013. Temporal patterns of diversification across global cichlid biodiversity (Acanthomorpha: Cichlidae). PLoS ONE 8: e71162.

Near, T.J., Dornburg, A., Eytan, R.I., Keck, B.P., Smith, W.L., Kuhn, K.L, Moore, J.A., Price, S.A., Burbrink, F.T., Friedman, M., and Wainwright, P.C. 2013. Phylogeny and tempo of diversification in the superradiation of spiny-rayed fishes. Proceedings of the National Academy of Sciences 110:12738-12743.

Owens, H.L., Campbell, L.P., Dornak, L.L., Saupe, E.E., Barve, N., Soberon, J., Ingenloff, K., Lira-Noriega, A., Hensz, C.M., Myers, C.E., and Peterson, A.T. 2013. Constraints on interpretation of ecological niche models by limited environmental ranges on calibration areas. Ecological Modeling 263: 10-18.

Schultze, H.-P., and  Arratia, G. 2013. The caudal skeleton of basal teleosts, its conventions, and some of its major evolutionary novelties in a temporal dimension. Pages 187-246, In: Mesozoic Fishes 5–Global Diversity and Evolution. Arratia, G., Schultze, H.-P., and Wilson, M.V.H. (eds.). Verlag Dr. Friedrich Pfeil, München, Germany.

Tischlinger, H., and Arratia, G. 2013. Ultraviolet light as a tool of investigating Mesozoic fishes with a focus on the ichthyofauna of the Solnhofen Limestone. Pages 549-560, In: Mesozoic Fishes 5–Global Diversity and Evolution. Arratia, G., Schultze, H.-P., and Wilson, M.V.H. (eds.). Verlag Dr. Friedrich Pfeil, München, Germany.